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Caption: Mudskipper (Periophthalmus sp.) in mud. This amphibious fish inhabits estuaries and swamp habitats. It moves across land using its limb- like pectoral fins. It breathes on land using highly vascular sacks on its mouth and gills. Its skin contains cells that are filled with a watery fluid, which cushion its movements. Its protruding eyes are adapted for vision above the water. Photographed in Sabah, Malaysian Borneo.

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Keywords: amphibian, amphibious, animal, aquatic, biological, biology, bony fish, borneo, crawling, evolution, fauna, fins, fish, land, limbs, malaysia, malaysian borneo, mud, mud-flat, mudskipper, nature, one, periophtalmus sp., pisces, sabah, silt, single, south-east asia, southeast asia, walking, wildlife, zoological, zoology

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