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Thomas Hunt Morgan (1866-1945), US geneticist, in his study at Columbia University, New York City, USA. Morgan studied zoology at the State College of Kentucky. His work in genetics is remembered for the use of the fruit fly Drosophila melanogaster, and for establishing many ideas in chromosome theory. It was Morgan who proved that the chromosome was the carrier of heredity, and that the chromosome consists of genes. He also discovered crossing over (the exchange of genes between chromosomes), and helped produce the first chromosome map. He was awarded the Nobel Prize in Physiology or Medicine (1933) and the Copley Medal (1939). Photographed circa 1910.
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