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RICHARD BROOKS / SCIENCE PHOTO LIBRARY RICHARD BROOKS / SCIENCE PHOTO LIBRARY
Territorial dispute between rival male brown-marbled grouper fish (Epinephelus fuscoguttatus). Groupers are protogynous hermaphrodites, with all fish being female when they first attain sexual maturity. In the absence of a male, the fittest female turns into a male. Males control territories and harems of females, defending them against other males. The brown-marbled grouper inhabits reefs and shallow seas throughout the tropical Indo-Pacific region. It can reach over a metre in length, and it feeds on smaller fish, octopus and crustaceans. Filmed in Palau.
Model release not required. Property release not required.
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